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Formaciya Dyadohta geologichna formaciya z visokim vmistom skam yanilostej roztashovana u pusteli Gobi na pivdni Mongoliyi Datuyetsya piznoyu krejdoyu priblizno 75 71 miljon rokiv tomu Tipovoyu miscevistyu ye Bayanzag sho vidoma yak Palayuchi skeli Sered skam yanilostej viyavleno chislenni reshtki reptilij vklyuchayuchi dinozavriv i ssavciv DyadohtaKrayina MongoliyaLezhachij bikAlagteegdVisyachij bikBarun GojotKoordinati 44 08 19 pn sh 103 43 40 sh d 44 13861111113877200 pn sh 103 72777777780778763 sh d 44 13861111113877200 103 72777777780778763 Zmist 1 Istoriya rozkopok 2 Paleobiota 2 1 Flora 2 2 Amfibiyi 2 3 Krokodilomorfi 2 4 Yashirki 2 5 Ssavci 2 6 Pterozavri 2 7 Cherepahi 2 8 Dinozavri 2 8 1 Alvareszavri 2 8 2 Ankilozavridi 2 8 3 Ptahi 2 8 4 Ceratopsi 2 8 5 Dromeozavri 2 8 6 Gadrozavri 2 8 7 Galskaraptorini 2 8 8 Ornitomimozavri 2 8 9 Oviraptorozavri 2 8 10 Pahicefalozavri 2 8 11 Troodontidi 2 8 12 Tiranozavridi 2 8 13 Zauropodi 3 PrimitkiIstoriya rozkopok red Formaciya Dyadohta bula vpershe zadokumentovana ta doslidzhena pid chas paleontologichnih ekspedicij Amerikanskogo muzeyu prirodnoyi istoriyi u 1922 1925 rokah Ekspediciyi ocholyuvav Roj Chepmen Endryus u kompaniyi Voltera Villisa Grejndzhera Komanda provela masshtabni doslidzhennya v regioni Bayanzag yakij voni nazvali Palayuchimi skelyami oskilki na zahodi soncya vidkladennya cogo miscya mali harakternij chervonuvatij kolir Pomitni znahidki vklyuchali pershi vidomi skam yanilosti oviraptora protoceratopsa zaurornitoida i velociraptora pershi pidtverdzheni yajcya dinozavra chastkove gnizdo oviraptora a takozh vikopnih ssavciv Deyaki z nih buli korotko opisani Genri Ferfildom Osbornom protyagom ostannih rokiv ekspedicij U 1927 roci formaciya bula oficijno opisana Berki ta Morrisom 1 2 U 1963 roci mongolskij paleontolog Demberelin Dashzeveg povidomiv pro vidkrittya novoyi vikopnoyi lokalizaciyi formaciyi Dzhadohta u Tugriken Shire 3 U1960 1970 h rokah polsko mongolska ta radyansko mongolska paleontologichni ekspediciyi zibrali novi zrazki protoceratopsa ta velociraptora v comu misci sho zrobilo ci vidi dinozavriv zvichajnim yavishem u Tugriken Shiri 4 Sered najvidomishih vidkritiv zroblenih u Tugriken Shire bijcivski dinozavri protoceratops ta velociraptor sho vedut bij 5 6 U 1980 h rokah radyansko mongolska paleontologichna ekspediciya viyavila v pusteli Gobi kilka bagatih skam yanilostyami misc Sered cih dilyanok Udin Sajr yake bagate na skam yanilosti avimimiv z menshoyu kilkistyu skam yanilostej ssavciv ta inshih dinozavriv 4 U 1993 roci grupi spilnoyi mongolo amerikanskoyi ekspediciyi za pidtrimki Mongolskoyi akademiyi nauk i Amerikanskogo muzeyu prirodnoyi istoriyi vidkrili nove misce vikopnih reshtok formaciyi Dyadohta pid nazvoyu Ukhaa Tolgod sho oznachaye Korichnevi pagorbi Yak i v poperednih miscyah Ukhaa Tolgod viyaviv znachnu kilkist dobre zberezhenih skam yanilostej vklyuchayuchi visoki koncentraciyi ssavciv dinozavriv yashirok i yayec Perevazhna bilshist ekzemplyariv iz ciyeyi miscevosti zazvichaj traplyayutsya v majzhe povnomu zchlenuvanni Zagalom yaksho porivnyuvati z inshimi miscyami skam yanilostej mezozoyu riznomanitnist skam yanilostej v Ukhaa Tolgod ye nadzvichajno visokoyu 7 8 Paleobiota red Flora red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaRadicites R gobiensis Bayanzag Dvadcyat koreniv 9 Traheofit jmovirno hvojne derevo Amfibiyi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaGobiates Neviznachenij Udin Sajr Chastkovij skelet z chastkovim urostilem 10 zhabaKrokodilomorfi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaArtzosuchus A brachicephalus Udin Sajr Chastkovij cherep 11 Krokodilomorf Gobiosuchus G kielanae Bayanzag Kilka zrazkiv iz chastkovimi cherepami ta skeletami 12 13 Gobizuhid nbsp G parvus Udin Sajr Chastkovij cherep i skelet 14 15 GobizuhidShamosuchus S djadochtaensis Bayanzag Uhaa Tolgod Dva cherepa i chastkovo skelet 16 17 ParaligatorYashirki red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAdamisaurus A magnidentatus Bayanzag Tugriken Shireh Ukhaa Tolgod Cherepi ta skeleti z kilkoh ekzemplyariv 18 19 Teyida Takozh prisutnij u formaciyi Barun Gojot I Bayan Mandahu Aiolosaurus A oriens Ukhaa Tolgod Nepovnij cherep i chastkovij skelet 19 Varanoyid Carusia C intermedia Bayanzag Ukhaa Tolgod Cherepi kilkoh ekzemplyariv 19 Karusiyidi Takozh prisutnij u formaciyi Barun Gojot i Bayan Mandahu Cherminotus C longifrons Tugriken Shireh Ukhaa Tolgod Cherepi ta chastkovij skelet 19 Varanoyid Takozh prisutnij u formaciyi Barun GojotCtenomastax C parva Zos Nepovnij cherep 19 Iguana Takozh prisutnij u formaciyi Barun GojotDzhadochtosaurus D giganteus Tugriken Shireh Chastkovij cherep 20 Makrocefalozavr Eoxanta E lacertifrons Ukhaa Tolgod Nepovnij cherep 19 Scinkomorf Takozh prisutnij u formaciyi Barun GojotEstesia E mongoliensis Bayanzag Ukhaa Tolgod Chastkovi cherepi ta zubi 19 Monsterzavr Takozh prisutnij u formaciyi Barun GojotFlaviagama F dzerzhinskii Tugriken Shireh Cherep i dva hrebci 21 Pristiagama Globaura G venusta Bayanzag Ukhaa Tolgod Chastkovi cherepi 19 Scinkopodibni Takozh prisutnij u formaciyi Barun GojotGobiderma G pulchrum Udyn Sayr Ukhaa Tolgod Cherepi ta vidbitki shkiri 19 Monsterzavr Takozh prisutnij u formaciyi Barun GojotGobinatus G arenosus Ukhaa Tolgod Chastkovij cherep 19 Teyida Takozh prisutnij u formaciyi Barun GojotHymenosaurus H clarki Ukhaa Tolgod Chastkovij cherep 19 Scinkopodibni Isodontosaurus I gracilis Bayanzag Tugriken Shireh Ukhaa Tolgod Zos Bagato cherepiv i chastkovij skelet 19 Iguana Macrocephalosaurus Indeterminate Ukhaa Tolgod Chastkovij cherep i skelet 19 Teyida Takozh prisutnij u formaciyi Barun GojotMimeosaurus M crassus Bayanzag Ukhaa Tolgod Zos Wash Chastkovij cherep 19 Akrodont Myrmecodaptria M microphagosa Ukhaa Tolgod Odin cherep 19 GekkonOvoo O gurval Little Ukhaa Tolgod Chastkovij cherep 22 Varan Parmeosaurus P scutatus Ukhaa Tolgod Zchlenovanij cherep i skelet 19 Scinkomorf Phrynosomimus P asper Ukhaa Tolgod Dva chastkovi cherepa 19 Akrodont Takozh prisutnij u formaciyi Barun GojotPriscagama P gobiensis Bayanzag Ukhaa Tolgod Nepovni cherepi 19 Priskagamida Takozh prisutnij u formaciyi Barun GojotSaichangurvel S davidsoni Ukhaa Tolgod Povnij cherep i skelet v artikulyaciyi 23 Iguanida Slavoia S darevskii Ukhaa Tolgod Cherepi i skelet 19 Scinkomorf Takozh prisutnij u formaciyi Barun GojotTelmasaurus T grangeri Bayanzag Chastkovij cherep i skelet 24 19 Varan Temujinia T ellisoni Tugriken Shireh Ukhaa Tolgod Kilka chastkovih cherepiv 19 Iguana Tchingisaurus T multivagus Ukhaa Tolgod Chastkovij cherep 19 Teyida nenazvanij scinkopodibnij Indeterminate Ukhaa Tolgod Chastkovij cherep 19 Scinkomorf Varanoidea indet Indeterminate Ukhaa Tolgod Chastkova verhnya shelepa i hrebec 19 Varanoyid Zapsosaurus Z sceliphros Tugriken Shireh Dva chastkovi cherepi 19 Iguana Ssavci red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAsiatherium A reshetovi Udyn Sayr Zchlenovanij cherep i skelet 25 Metateriyi nbsp Bulganbaatar B nemegtbaataroides Bayanzag Ukhaa Tolgod Chastina cherepa ta inshi reshtki 7 Bagagorbkozubi Catopsbaatar C catopsaloides Ukhaa Tolgod Dyadohtateriyidi Takozh prisutnij u formaciyi Barun Gojot nbsp Chulsanbaatar C vulgaris Ukhaa Tolgod Cherep i chastkovij skelet 7 Bagagorbkozubi Takozh prisutnij u formaciyi Barun GojotDeltatheridium D pretrituberculare Bayanzag Ukhaa Tolgod Chastkovo zalishki cherepa ta skeleta 26 27 Tribosfenida Deltatheroides D cretacicus Bayanzag Chastkovij cherep 26 DyadohtateriyidiDjadochtatherium D matthewi Bayanzag Tugriken Shireh Chastkovi cherepi 28 29 DyadohtateriyidiHyotheridium H dobsoni Bayanzag Chastkovij cherep 26 Teriyi Indeterminate Ukhaa Tolgod Teriyi Kamptobaatar K kuczynskii Bayanzag Ukhaa Tolgod Chastina cherepa ta inshi reshtki 30 7 BagagorbkozubiKennalestes K gobiensis Bayanzag Ukhaa Tolgod Majzhe povnij cherep ta inshi ostanki 31 7 Vishi zviri Kryptobaatar K dashzevegi Bayanzag Tugriken Shireh Ukhaa Tolgod Cherepi ta zalishki skeletiv kilkoh ekzemplyariv 30 32 DyadohtateriyidiMaelestes M gobiensis Ukhaa Tolgod Chastkova chastina cherepa zi skeletom 33 Cimolesti Mangasbaatar M udanii Udyn Sayr Cherepi ta chastkovij skelet dvoh ekzemplyariv 34 DyadohtateriyidiNemegtbaatar N gobiensis Ukhaa Tolgod Bagagorbkozubi Takozh prisutnij u formaciyi Barun Gojot nbsp Sloanbaatar S mirabilis Bayanzag Ukhaa Tolgod Povnij cherep ta inshi reshtki 30 7 BagagorbkozubiTombaatar T sabuli Ukhaa Tolgod Chastkovij cherep 35 DyadohtateriyidiUkhaatherium U nessovi Ukhaa Tolgod Chastkovi abo majzhe povni skeleti kilkoh osobin 36 37 38 Vishi zviri Zalambdalestes Z lechei Bayanzag Tugriken Shireh Cherepi ta skeleti kilkoh ekzemplyariv 26 39 Vishi zviri nbsp Pterozavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAzhdarchidae indet Neviznachenij Tugriken Shire Neviznachena kistka v kishkovij porozhnini velociraptora 40 AzhdarhidCherepahi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaNanhsiungchelyidae indet Neviznachenij Abdrant Nuru Tri oskolki pancira 41 Nanhsiungchelyid Neviznachenij Bayanzag Chastkovi panciri 41 Nanhsiungchelyid Neviznachenij Udin Sajr Dva oskolki pancira 41 Nanhsiungchelyid Zangerlia Z dzamynchondi Zamin Hondt Chastkova obolonka 42 Nanhsiungchelyid Z ukhaachelys Uhaa Tolgod Chastkovij cherep i skelet 43 Nanhsiungchelyid Dinozavri red Alvareszavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAlvarezsauridae indet Neviznachenij Tugriken Shire Chastkovo elementi cherepa mozkovoyi korobki ta skeleta dvoh zrazkiv 44 45 Kol K ghuva Uhaa Tolgod Prava noga 46 nbsp Shuvuuia S deserti Uhaa Tolgod Kilka ekzemplyariv iz cherepom i skeletami 47 48 49 50 51 nbsp Neopisani Alvarezsauridae Neviznachenij Bayanzag Chastkovij tazovij poyas i zadnya kincivka Neviznachenij Gilvent WashAnkilozavridi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaMinotaurasaurus M ramachandrani Uhaa Tolgod Dva povnih cherepa nizhnya shelepa ta pershe shijne pivkilce 52 53 Ankilozavrid yakij ranishe vvazhavsya molodshim sinonimom Tarchia ale teper vvazhayetsya dijsnim i okremim taksonom nbsp Pinacosaurus P grangeri Bayanzag Uhaa Tolgod Tri cherepa nizhnya shelepa peredzubna kistka shijni hrebci spinni hrebci hvostovi hrebci rebra lopatka korakoyidi plechova kistka promeneva kistka liktova kistka klubova kistka stegnova kistka velikogomilkova kistka malogomilkova kistka taz manus ruchki hvostovoyi bulavi shijni pivkilcya osteodermi ta majzhe povnij skelet bez cherepa 54 55 56 Ankilozavrid takozh vidomij iz formaciyi Alagteyeg i formaciyi Bayan Mandahu nbsp Ankylosauridae indet Neviznachenij Zamin Hondt Chastkovij postkranialnij skelet z in situ osteodermami 57 Ranishe nazivavsya Saichania ale teper zgaduyetsya yak Ankylosauridae indet abo por Pinakozavr 57 nbsp Ptahi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaApsaravis A ukhaana Uhaa Tolgod Chastkovij postkranialnij skelet 58 Bazalnij orniturovij ptah Elsornis E keni Tugriken Shire Chastkovo zchlenovanij skelet bez cherepa 59 EnanciornisGobipteryx G minuta Uhaa Tolgod Chastkovij cherep 60 Enantiornis Takozh prisutnij u formaciyi Barun Goyot Protoceratopsidovum P fluxuosum Bayanzag Fragmenti yayec 61 Yajcya jmovirno zneseni ptahom 62 P minimum Baga Tariah Tugriken Shire Kladka z chotiroh yayec i odniyeyi kladki yajcya 61 Yajcya jmovirno zneseni ptahom 62 P sincerum Bayanzag Tugriken Shire Kilka yayec i shkaralupi 61 Yajcya jmovirno zneseni ptahom 62 Styloolithus S sabathi Bayanzag Yajcya 62 Yajcya jmovirno zneseni ptahom Ceratopsi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaBainoceratops B efremovi Bayanzag Chastkovi hrebci 63 Protoceratopsid Mozhe buti sinonimom Protoceratops 64 Bagaceratops Neviznachenij Udin Sajr Cherep iz chastkovim skeletom 65 Protoceratopsid Neviznachenij mizh Bagaceratops i Protoceratops 65 nbsp Protoceratops P andrewsi Bayanzag Tugriken Shire Udin Sajr Zamin Hondt Kilka chastkovih ta povnih zrazkiv 66 67 68 69 65 Protoceratopsid nbsp P hellenikorhinus Bor Tolgoj Udin Sajr Chastkovi reshtki cherepa 70 Protoceratopsid nbsp Protoceratopsidae indet Neviznachenij Uhaa Tolgod Chislenni cherepi ta ostanki 71 72 Protoceratopsid nbsp Udanoceratops U tschizhovi Udin Sajr Cherep i fragmentovani elementi skeleta 73 Gigantskij leptoceratopsid nbsp Dromeozavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaTsaagan T mangas Uhaa Tolgod Cherep i chastkovij skelet 74 nbsp Velociraptor V mongoliensis Bayn Dzak Chimney Buttes Gilvent Wash Tugriken Shireh Udyn Sayr Ukhaa Tolgod Kilka chastkovih ta povnih zrazkiv 75 76 77 78 79 80 nbsp Neopisani Dromaeosauridae Neviznachenij Abdrant Nuru Kigot 81 Neviznachenij Zos Uosh Lobova oblast 74 Vidriznyayetsya vid Caagan 74 Gadrozavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaHadrosauroidea indet Neviznachenij Tugriken Shire Fragmentovani reshtkinepovnolitnih 82 83 Plesiohadros P djadokhtaensis Alag Teg Cherep i chastkovi elementi tila 83 Galskaraptorini red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaHalszkaraptor H escuilliei Uhaa Tolgod Chastkovij skelet iz povnim cherepom 84 nbsp Mahakala M omnogovae Tugriken Shire Fragmentovanij cherep i skelet 85 nbsp Ornitomimozavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAepyornithomimus A tugrikinensis Tugriken Shire Majzhe povna stopa 86 Ornitomimid nbsp Ornithomimosauria indet Neviznachenij Uhaa Tolgod Chastkova mozkova obolonka kinchiki shelep rebra ta fragmenti hrebciv 87 88 OrnitomimidOviraptorozavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAvimimus A portentonsus Udin Sajr Chastkovij cherep i skelet 89 90 Avimimid Jogo misceve pohodzhennya bulo spirnim i mozhlivo pohodit vid formaciyi Nemegt 91 nbsp Citipati C osmolskae Uhaa Tolgod Kilka zrazkiv iz chastkovimi chi majzhe povnimi skeletami embrionami yajcyami ta gnizdovimi osobinami 92 93 94 95 Oviraptorid nbsp Elongatoolithus E frustrabilisE subtitectoriusKhaan K mckennai Uhaa Tolgod Kilka zrazkiv iz chastkovimi abo povnimi skeletami ta cherepami 93 96 97 Oviraptorid nbsp Macroolithus M mutabilis Yajcya Yajcya jmovirno vidkladeni oviraptoridomOviraptor O philoceratops Bayanzag Chastkovij skelet iz cherepom pov yazanij iz gnizdom i molodnyakom 98 94 95 Oviraptorid nbsp Oviraptoridae indet Neviznachenij Zamin Hondt Majzhe povnij skelet iz cherepom 99 Oviraptorid takozh vidomij yak oviraptorid Zamyn Khondt Nepevne posilannya na Citipati 93 94 nbsp Neviznachenij Zamin Hondt Majzhe povnij cherep z atlasom i vissyu Oviraptorid nbsp Neviznachenij Udin Sajr Zibrannya osib 95 OviraptoridNeviznachenij Ne viznacheno Dva cherepa z harakternim visokim grebenem 100 101 Oviraptorid nbsp Pahicefalozavri red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaGoyocephale G lattimorei Boro Hovil Chastkovij cherep i skelet 102 Pahicefalozavrid Do ciyeyi formaciyi mozhut nalezhati vidkladennya miscevosti 102 103 nbsp Troodontidi red Rid Vidi Misceznahodzhennya Material Primitki ZobrazhennyaAlmas A ukhaa Uhaa Tolgod Cherep iz chastkovim skeletom 104 nbsp Archaeornithoides A deinosauriscus Bayanzag Chastkovij cherep 105 Troodontid Neviznacheni vidnosini mizh celurozavrami 105 106 Byronosaurus B jaffei Uhaa Tolgod Cherep i oskolkovij skelet 107 nbsp Gobivenator G mongoliensis Zamin Hondt Majzhe povnij skelet 108 nbsp Saurornithoides S mongoliensis Bayanzag Cherep z oskolkovim skeletom 109 nbsp Troodontidae indet Neviznachenij Uhaa Tolgod Chastkovij skelet 110 Neviznachenij Uhaa Tolgod Molodi cherepa skelet i odne gnizdo 106 Troodontid Vidnositsya do Almasa 111 104 abo Bajronozavra 106 nbsp Neviznachenij Uhaa Tolgod Fragmentovanij cherep i zalishki skeleta 112 Troodontid Poperedno vidnesenij do Saurornithoides 112 ale teper viklyuchenij 109 Neviznachenij Uhaa Tolgod Chastkovij cherep i skeleti dvoh ekzemplyariv 48 Tiranozavridi red Rid Vidi Misceznahodzhennya 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